Ecology

I

Introduction

Ecology, the study of the relationship of plants and animals to their physical and biological environment. The physical environment includes light and heat or solar radiation, moisture, wind, oxygen, carbon dioxide, nutrients in soil, water, and atmosphere. The biological environment includes organisms of the same kind as well as other plants and animals.

Because of the diverse approaches required to study organisms in their environment, ecology draws upon such fields as climatology, hydrology, oceanography, physics, chemistry, geology, and soil analysis. To study the relationships between organisms, ecology also involves such disparate sciences as animal behavior, taxonomy, physiology, and mathematics.

An increased public awareness of environmental problems has made ecology a common but often misused word. It is confused with environmental programs and environmental science (see Environment). Although the field is a distinct scientific discipline, ecology does indeed contribute to the study and understanding of environmental problems.

The term ecology was introduced by the German biologist Ernst Heinrich Haeckel in 1866; it is derived from the Greek oikos (“household”), sharing the same root word as economics. Thus, the term implies the study of the economy of nature. Modern ecology, in part, began with Charles Darwin. In developing his theory of evolution, Darwin stressed the adaptation of organisms to their environment through natural selection. Also making important contributions were plant geographers, such as Alexander von Humboldt, who were deeply interested in the “how” and “why” of vegetational distribution around the world.

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II

The Earth's Biosphere

The thin mantle of life that covers the earth is called the biosphere. Several approaches are used to classify its regions.

A

Biomes

The broad units of vegetation are called plant formations by European ecologists and biomes by North American ecologists. The major difference between the two terms is that biomes include associated animal life. Major biomes, however, go by the name of the dominant forms of plant life.

Influenced by latitude, elevation, and associated moisture and temperature regimes, terrestrial biomes vary geographically from the tropics through the arctic and include various types of forest, grassland, shrub land, and desert. These biomes also include their associated freshwater communities: streams, lakes, ponds, and wetlands. Marine environments, also considered biomes by some ecologists, comprise the open ocean, littoral (shallow water) regions, benthic (bottom) regions, rocky shores, sandy shores, estuaries, and associated tidal marshes.

See also Chaparral; Coral Reef; Estuary; Marine Life; Marshland; Peatland; Savanna; Shore Life; Tundra.

B

Ecosystems

A more useful way of looking at the terrestrial and aquatic landscapes is to view them as ecosystems, a word coined in 1935 by the British plant ecologist Sir Arthur George Tansley to stress the concept of each locale or habitat as an integrated whole. A system is a collection of interdependent parts that function as a unit and involve inputs and outputs. The major parts of an ecosystem are the producers (green plants), the consumers (herbivores and carnivores), the decomposers (fungi and bacteria), and the nonliving, or abiotic, component, consisting of dead organic matter and nutrients in the soil and water. Inputs into the ecosystem are solar energy, water, oxygen, carbon dioxide, nitrogen, and other elements and compounds. Outputs from the ecosystem include water, oxygen, carbon dioxide, nutrient losses, and the heat released in cellular respiration, or heat of respiration. The major driving force is solar energy.